Plants 10-40 cm; self-incom-patible. Stems (± reddish distally) simple or branched (branches 1-4+ and ascending, or more numerous and spreading, rigid), hirsutulous-hispidulous, ± villous. Leaves (sometimes basal clusters persisting to flowering) mostly alternate, 2-5 cm, ± hispidulous (margins ± densely ciliate, especially proximally). Heads in ± spi-ciform arrays (1-3 per node). Peduncular bracts (crowded, closely investing, obscuring heads before flowering) lance-elliptic to linear (± flat distally), 3-10(-15) mm (hirsutulous, bristly and/or pectinate-fimbriate, deeply grooved proximally), apices (often ± reddish) rounded to truncate, tack-glands (0-)1 (terminal). Phyllaries (± reddish distally) 5-6 mm, abaxial faces hirsutulous, usually bristly, ± shaggy long-hairy near distal margins, tack-glands 0. Paleae 5-6 mm. Ray florets 1-4; corollas white to pinkish, tubes ca. 2 mm (papillate), laminae ca. 5 mm (central lobes much narrower than laterals, widest at bases, symmetric, laterals asymmetric, sinuses 1/3-1/2 laminae). Disc florets 5-15; corollas white to pinkish, 5-6 mm. Ray cypselae ca. 2.5 mm, smooth, densely appressed-hairy. Disc cypselae ca. 2.5 mm, appressed-hairy, especially on angles; pappi of ca. 10 lanceolate-aristate scales ca. 4-6 mm. 2n = 14. Flowering spring-summer. Dry, rocky hills, ridges; 300-1100 m; Calif. Calycadenia villosa occurs in Monterey and San Luis Obispo counties and is often localized. It is in the Center for Plant Conservation´s National Collection of Endangered Plants.