Leaf blade lanceolate to oblong, 3--4-pinnate, proximal pinnae 1/4--1/3 as long as blades. n = 2 n = 81, apogamous. Sporulating summer--fall. Rocky slopes and cliffs; usually on calcareous or volcanic substrates; 1700--2500 m; Ariz., N.Mex., Tex.; n Mexico.