Sporangia containing 32 spores; spores averaging 50--65 µm diam. n = 2 n = 87, apogamous. Sporulating summer--fall. Rocky slopes and cliffs; occurring on calcareous and noncalcareous substrates; 800--2100 m; Ariz., Ga., N.Mex., Tex.; Mexico; West Indies; Central America; South America.
