Biennials or perennials; caudex unbranched or with a root crown, often slender. Basal leaves: (petiole often hirsute); blade margins lyrate or lyrate-pinnatifid, terminal lobes distinct, (surfaces pubescent). Petals 6-8 mm. Fruits 0.8-1 mm wide; style 0.5-1 mm. 2n = 16, 32. Flowering Apr-Jul. Cliffs, ledges, thickets, stream and river banks, woods, limestone crevices and bluffs, sandstone hills and outcrops, serpentine rocks and barrens, shale, talus, sand dunes; 0-1000 m; Alta., B.C., Man., N.W.T., Ont., Sask.; Alaska, Conn., Del., D.C., Ill., Ind., Iowa, Md., Mass., Mich., Minn., Miss., Mo., N.J., N.Y., N.C., Ohio, Pa., Tenn., Vt., Va., W.Va., Wis.