Annuals or biennials [perennials]; not scapose; pubescent, trichomes stellate, mixed with simple ones. Stems erect [ascending], usually branched distally. Leaves basal and cauline; petiolate or sessile; basal not rosulate, petiolate, blade margins entire or repand [dentate, sinuate]; cauline (middle and distal) sessile. Racemes (corymbose, several-flowered, dense), considerably elongated in fruit. Fruiting pedicels erect or divaricate, slender. Flowers: sepals erect-ascending [suberect, spreading], oblong, lateral pair not saccate basally; petals usually white, rarely yellow, obcordate, apex deeply 2-fid; stamens tetradynamous; filaments: median pair flattened basally, unappendaged, [laterally 1-toothed], lateral pair with basal toothlike appendage; anthers oblong, (apex obtuse); nectar glands (4), lateral, 1 on each side of lateral stamen. Fruits sessile, oblong, or elliptic [ovate, obovate, or orbicular], smooth, slightly inflated [or not inflated], latiseptate; valves each not veined or with obscure midvein, stellate-hairy [glabrous]; replum rounded; septum complete, (membranous); ovules 4-16 per ovary; stigma capitate, obscurely 2-lobed. Seeds biseriate, flattened [plump], margined [winged or not], lenticular or ovoid-lenticular [suborbicular]; seed coat (minutely reticulate), not mucilaginous when wetted; cotyledons accumbent. x = 8.