Perennials, subshrubs, or shrubs [trees, vines], 40-200+ cm. Stems erect, branched. Leaves cauline; mostly opposite (distal sometimes alternate); petiolate; blades usually 3-nerved from bases, mostly deltate or ovate to triangular-lanceolate or lanceolate (lacking internal translucent lenses or streaks), margins entire or dentate, faces glabrous or sparsely hispidulous, puberulent, or strigose, sometimes sparsely to densely gland-dotted. Heads discoid, in corymbiform to paniculiform arrays. Involucres obconic to hemispheric, 2-3 mm diam. Phyllaries persistent (at least outermost), 7-16 in 1-2[-4] series, 3-nerved, ovate to lanceolate, ± equal or unequal. Receptacles flat or convex, epaleate. Florets 3-13[-20]; corollas whitish to pinkish [violet], throats funnelform (lengths ca. 2.5 times diams.); styles: bases sometimes enlarged, glabrous, branches filiform to obscurely clavate. Cypselae prismatic, 5-ribbed, sparsely hispidulous or piloso-strigose, sometimes gland-dotted; pappi persistent, of 30-35, barbellate bristles in 1 series. x = 10.