Perennials, 20-150 cm. Stems erect, usually branched. Leaves basal and cauline; opposite (proximal) and alternate; usually petiolate (at least proximal); blades (usually 3- or 5-nerved from at or near bases) elliptic, lanceolate, lance-linear, linear, oblanceolate, ovate, ovate-lanceolate, or spatulate, bases mostly cuneate, margins entire, faces glabrous or hairy. Heads radiate, borne singly or (2-15+) in corymbiform arrays. Involucres campanulate to hemispheric or broader, 10-50 mm diam. Phyllaries persistent, 22-32+ in ± 3 series (mostly deltate-ovate or ovate to lanceolate, linear, or oblong, subequal or unequal, outer sometimes foliaceous). Receptacles ± convex, paleate (paleae falling with cypselae, conduplicate, scarious or chartaceous). Ray florets 8-21, neuter; corollas ± yellow. Disc florets 30-200+, bisexual, fertile; corollas yellow or purple to brown, tubes shorter than or equaling throats, lobes 5, deltate (abaxially hairy). Cypselae (brownish) ± compressed, obovate to obcordate (margins usually ± winged, usually ± ciliate, faces glabrous or strigose); pappi 0, or persistent, of 2 ± subulate scales plus 0-4+ shorter, lacerate scales (sometimes all ± connate). x = 15.