Perennials, 5-60(-100) cm (taproots relatively massive; caudices sometimes branched). Stems erect and branched (mostly distally), or decumbent (and seldom branched distally). Leaves basal and cauline (basal usually scalelike on flowering stems, well-formed on non-flowering shoots); alternate; petiolate; blades (3-or 5-nerved or pinnately nerved), cordate-ovate, deltate, deltate-triangular, ovate, ovate-lanceolate, or suborbiculate, bases broadly cuneate, subcordate, or truncate, margins entire or dentate to crenate, faces glabrous or hairy (often gland-dotted). Heads radiate or discoid, borne singly (terminal) or in ± racemiform arrays. Involucres campanulate to ± hemispheric, 12-40 mm diam. Phyllaries persistent, (8-)12-26+ in 2-3+ series (subequal to unequal, outer usually surpassing inner). Receptacles flat to convex, paleate (paleae persistent, conduplicate, chartaceous). Ray florets 0 or (1-)5-21+, pistillate, fertile; corollas yellow. Disc florets 50-150+, bisexual, fertile; corollas yellow, tubes shorter than cylindric throats, lobes 5, lance-deltate (style branches stigmatic in 2, barely distinct lines, appendages filiform). Cypselae ± prismatic, 3-4-angled (faces glabrous or strigillose); pappi usually persistent, coroniform (± lacerate, sometimes with 1-4 teeth or scales), sometimes 0. x = 19.