Plants not scapose; pubescent, trichomes sessile, medifixed, appressed, 2-rayed (malpighiaceous) or 3-5(-8)-rayed (stellate), rays (when 2) parallel to long axis of stems, leaves, sepals, and fruits. Stems erect or ascending [decumbent], unbranched or branched basally and/or distally. Leaves basal and cauline; petiolate or sessile; basal rosulate or not, petiolate, blade margins usually entire, dentate, sinuate-dentate, or denticulate, rarely pinnatifid or pinnatisect; cauline petiolate or sessile, blade (base cuneate or attenuate [auriculate]), margins entire, dentate, denticulate, dentate-sinuate, or repand. Racemes (densely flowered, E. pallasii bracteate basally). Fruiting pedicels erect, ascending, divaricate, reflexed, horizontal, or spreading, slender or stout (nearly as wide as fruit). Flowers: sepals oblong or linear, lateral pair saccate or not basally (pubescent); petals suborbicular, obovate, or spatulate, claw differentiated from blade (subequaling or longer than sepals, apex rounded [emarginate]); stamens (erect), tetradynamous; filaments not dilated basally; anthers oblong or linear; nectar glands (1, 2, or 4), distinct or confluent, subtending bases of stamens, median glands present or absent. Fruits usually sessile, rarely shortly stipitate (gynophore to 4 mm), usually linear or narrowly so [oblong], smooth or torulose, (keeled or not); valves each with obscure to prominent midvein, pubescent outside, usually glabrous inside; replum rounded; septum complete, (not veined); ovules [15-]20-120 per ovary; (style relatively short, rarely 1/2 as long as or subequaling fruit, often pubescent); stigma capitate. Seeds plump or flattened, winged, margined, or not winged, oblong, ovoid, obovate, or suborbicular; seed coat (minutely reticulate), mucilaginous when wetted; cotyledons incumbent, rarely accumbent. x = (6) 7, 8 (9-17).