Plant: perennial herb; 3-6 dm high, nearly glabrous, erect Leaves: 4 per node, 13-31 mm long, linear to broadly lanceolate, 3-veined, minutely scaberulous, the apices mostly obtuse INFLORESCENCE: flowers perfect, numerous in a more or less condensed terminal, nearly leafless panicle of cymules Flowers: corollas rotate or a little cupped at base, the 4 lobes ovate, the apices blunt; ovaries generally densely hairy with short upwardly-curved hairs Fruit: FRUITS dry, with short bristles, the mericarps becoming reniform Misc: Meadows, streamsides, moist shady places; 2150-2900 m (7000-9500 ft); Jun-Aug REFERENCES: Dempster, Lauramay T. 1995. Rubiaceae. J. Ariz. – Nev. Sci. 29(l): 29.